[1] Univ Oulu, Ecol & Genet Res Unit, Oulu, Finland
[2] UiT Arctic Univ Norway, Ctr Arctic Gas Hydrate Environm & Climate, Dept Geosci, Tromso, Norway
[3] Univ Alaska Anchorage, Dept Geol Sci, Anchorage, AK USA
[4] Natl Ctr Sci Res, Inst Geosci & Environm, Grenoble, France
[5] Univ Oulu, Water Energy & Environm Engn Res Unit, Oulu, Finland
[6] Univ Alaska Anchorage, Dept Biol Sci, Anchorage, AK USA