[1] Ifremer Ctr Bretagne, Lab Environm Profond, F-29280 Plouzane, France
[2] Univ Georgia, Dept Marine Sci, Athens, GA 30602 USA
[3] Univ Leeds, Sch Earth & Environm, Leeds LS2 9JT, W Yorkshire, England
[4] Ifremer Ctr Bretagne, Unite Geosci Marines, F-29280 Plouzane, France
[5] Leibniz Inst Baltic Sea Res Warnemuende IOW, Dept Phys Oceanog & Instrumentat, Seestr 15, D-18119 Warnemunde, Germany