[1] Univ Michigan, Dept Mol & Integrat Physiol, Ann Arbor, MI 48109 USA
[2] Univ Michigan, Dept Med, Ann Arbor, MI 48109 USA
[3] VA Ann Arbor Healthcare Syst, Ann Arbor, MI USA
[4] Columbia Univ Coll Phys & Surg, Dept Pathol & Cell Biol, Taub Inst Res Alzheimers Dis & Aging Brain, 630 W 168th St, New York, NY 10032 USA