[1] Univ Sheffield, Dept Anim & Plant Sci, Sheffield S10 2TN, S Yorkshire, England
[2] Univ British Columbia, Dept Forest Sci, Vancouver, BC V6T 1Z4, Canada
[3] Univ Montpellier 2, ISEM, F-34095 Montpellier 05, France
[4] Univ E Anglia, Climat Res Unit, Climate Impacts LINK Project, Norwich NR4 7TJ, Norfolk, England