[1] Univ Sydney, Sch Biol Sci A08, Sch Life & Environm Sci, Sydney, NSW 2006, Australia
[2] Univ Haifa, Fac Nat Sci, Dept Evolutionary & Environm Biol, Inst Evolut, IL-3498838 Haifa, Israel
[3] Univ Tennessee, Dept Ecol & Evolut Biol, Knoxville, TN 37996 USA