[1] Hong Kong Univ Sci & Technol, Div Life Sci, Hong Kong, Hong Kong, Peoples R China
[2] China Geol Survey, Guangzhou Marine Geol Survey, MLR Key Lab Marine Mineral Resources, Guangzhou, Guangdong, Peoples R China
[3] Chinese Acad Sci, South China Sea Inst Oceanol, CAS Key Lab Ocean & Marginal Sea Geol, Guangzhou, Guangdong, Peoples R China
[4] Hong Kong Baptist Univ, Dept Biol, Hong Kong, Hong Kong, Peoples R China