[1] Univ Tasmania, Dairy Ctr, Tasmanian Inst Agr, Burnie, Tas 7320, Australia
[2] Dept Environm & Primary Ind, Div Agr Res, Ellinbank, Vic 3821, Australia
[3] Univ Melbourne, Melbourne Sch Land & Environm, Parkville, Vic 3010, Australia
[4] Univ Sydney, Fac Vet Sci, MC Franklin Lab, Camden, NSW 2570, Australia
[5] DairyNZ, Hamilton 3240, New Zealand