[1] Univ Guelph, Dept Anim & Poultry Sci, Guelph, ON N1G 2W1, Canada
[2] Eli Lilly & Co, Elanco Anim Hlth Provel Div, Res Pk Ctr, Guelph, ON N1G 4T2, Canada
[3] Univ Guelph, Dept Populat Med, Guelph, ON N1G 2W1, Canada
[4] Univ Manitoba, Dept Anim Sci, Winnipeg, MB R3T 2N2, Canada