[1] Tokyo Univ Agr & Technol, Res & Educ Ctr Prevent Global Infect Dis Anim, 3-5-8 Saiwai, Fuchu, Tokyo 1838509, Japan
[2] Gifu Univ, United Grad Sch Vet Sci, 1-1 Yanagito, Gifu 5011193, Japan
[3] Gifu Cent Livestock Hyg Serv Ctr, 4-2-22 Imarnine, Gifu 5008388, Japan
[4] Tokyo Univ Agr & Technol, Inst Agr, 3-5-8 Saiwai, Fuchu, Tokyo 1838509, Japan
[5] Azabu Univ, Sch Vet Med, Chuo Ku, 1-17-71 Fuchinobe, Sagamihara, Kanagawa 2525201, Japan
[6] Ishikawa Prefectural Univ, Fac Bioresources & Environm Sci, 1-308 Suematsu, Nonoichi, Ishikawa 9218836, Japan