[1] INRA, UMR AMAP, F-34398 Montpellier 5, France
[2] Pousse Conseil, F-34970 Lattes, France
[3] Scottish Crop Res Inst, Dundee DD2 5DA, Scotland
[4] CIRAD, UMR AMAP, F-34398 Montpellier 5, France
[5] Appalachian State Univ, Dept Geol, Boone, NC 28608 USA