[1] Univ Fed Ceara, Ctr Ciencias, Dept Biol, Fortaleza, Ceara, Brazil
[2] Inst Pesquisas Jardim Bot Rio de Janeiro, Escola Nacl Bot Trop, Programa Posgrad Bot, Rio De Janeiro, Brazil
[3] Univ Fed Pernambuco, Dept Nucl Energy, Recife, PE, Brazil
[4] Univ Fed Rural Pernambuco, Dept Biol, Recife, PE, Brazil