[1] Kyushu Univ, Fac Agr, Dept Biosci & Biotechnol, Nishi Ku, 744 Motooka, Fukuoka 8190395, Japan
[2] Tohoku Med & Pharmaceut Univ, Dept Infect & Host Def, Sendai, Miyagi 9818558, Japan
[3] Sojo Univ, Fac Biotechnol & Life Sci, Dept Appl Microbial Technol, Kumamoto 8600082, Japan