[1] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[2] Lab Nacl Identificat & Caracterizac Vegetal LaniV, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[3] Milwaukee Publ Museum, Bot Dept, 800 W Wells St, Milwaukee, WI 53233 USA
[4] UNAM, Inst Biol, Dept Bot, Ciudad Univ,Apartado Postal 70-367, Ciudad De Mexico 04510, Mexico
[5] Inst Ecol, Biol Evolut, AC Carretera Antigua Coatepec 351, Xalapa 91073, Veracruz, Mexico
[6] Univ Ciencias & Artes Chiapas, Fac Ciencias Biol, Herbario Eizi Matuda, Libramiento Norte Poniente 1150, Tuxtla Gutierrez 29039, Chiapas, Mexico