[1] Univ Estadual Regiao Tocantina Maranhao, Rua Godofredo Viana 1300, BR-65901480 Imperatriz, MA, Brazil
[2] Conjunto Jardim Guaruma 28, BR-67013880 Ananindeua, PA, Brazil
[3] Univ Fed Para, Progama Posgrad Ciencias Ambientais, Rua Augusto Correa 1, BR-66075110 Belem, Para, Brazil
[4] Univ Fed Rural Amazonia, Tv Pau Amarelo S-N, BR-68650000 Capitao Poco, PA, Brazil
[5] Inst Nacl Colonizacao & Reforma Agr, Rod Murucutum S-N, BR-66610903 Belem, Para, Brazil
[6] Univ Fed Rural Amazonia, Av Presidente Tancredo Neves 2501, BR-66077830 Belem, Para, Brazil