[1] INRA, DGER, UMR 1225, F-31076 Toulouse, France
[2] Moredun Res Inst, Penicuik EH26 OPZ, Midlothian, Scotland
[3] Scottish Agr Coll, Edinburgh EH9 3JG, Midlothian, Scotland
[4] Royal Vet & Agr Univ, DK-1870 Frederiksberg C, Denmark
[5] Massey Univ, Inst Vet Anim & Biomed Sci, Palmerston North, New Zealand