[1] Univ New England, Sch Rural Sci & Agr, Cooperat Res Ctr Beef Genet Technol, Armidale, NSW 2351, Australia
[2] Univ New England, Dept Primary Ind, Cooperat Res Ctr Beef Genet Technol, Armidale, NSW 2351, Australia
[3] Univ New England, Sheep Ind Cooperat Res Ctr, Cooperat Res Ctr Beef Genet Technol, Armidale, NSW 2351, Australia
[4] Murdoch Univ, Div Vet & Biomed Sci, Cooperat Res Ctr Beef Genet Technol, Murdoch, WA 6150, Australia
[5] Univ New England, Anim Genet & Breeding Unit, Cooperat Res Ctr Beef Genet Technol, Armidale, NSW 2351, Australia