[1] Univ Malaysia Terengganu, Fac Fisheries & Food Sci, Dept Fisheries & Aquaculture, Kuala Nerus, Terengganu, Malaysia
[2] Univ Malaysia Terengganu, Inst Marine Biotechnol, Kuala Nerus, Terengganu, Malaysia
[3] Natl Fish Hlth Res Div, Batu Maung, Penang, Malaysia
[4] Univ Putra Malaysia, Fac Sci, Dept Biol, Serdang, Selangor, Malaysia
[5] Univ Malaysia Terengganu, Inst Trop Aquaculture & Fisheries Res, Kuala Nerus, Terengganu, Malaysia