[1] Univ Fed Mato Grosso, Inst Ciencias Nat Humanas & Sociais, Programa Posgrad Ciencias Ambientais, Av Alexandre Ferronato 1-200, BR-78557267 Sinop, MT, Brazil
[2] Russian Acad Sci, Inst Problems Ecol & Evolut, Moscow, Russia
[3] Univ Fed Piaui, Ave Cicero Eduardo S-N, BR-64607675 Picos, PI, Brazil
[4] Univ Fed Mato Grosso, Inst Biociencias, Dept Biol & Zool, Ave Fernando Correa da Costa 2-367, BR-78060900 Cuiaba, MT, Brazil