Resonance in Beverton-Holt population models with periodic and random coefficients

An increase in the mean population density in a fluctuating environment is known as resonance. Resonance has been observed in laboratory experiments and has been studied in discrete-time population models. We investigate this phenomenon in the Beverton-Holt model with either periodic or random variables for two biologically relevant coefficients: the intrinsic growth rate and the carrying capacity. Three types of resonance are defined: arithmetic, geometric and harmonic. Conditions are derived for each type of resonance in the case of period-2 coefficients and some results for period p > 2. For period 2, regions in parameter space where each type of resonance occurs are shown to be subsets of each other. For the case of random coefficients with constant intrinsic growth rate, it is shown that the three types of resonance do not occur. Numerical examples illustrate resonance and attenuance (decrease in the mean population density) in the Beverton-Holt model when the coefficients are discrete random variables.