[1] Univ Toronto, Dept Ophthalmol & Vis Sci, Toronto, ON, Canada
[2] Univ Toronto, Dept Mol Genet, Toronto, ON, Canada
[3] Maisonneuve Rosemont Hosp, Ctr Rech, Montreal, PQ, Canada
[4] Toronto Western Res Inst, Vis Sci Res Program, Toronto, ON, Canada