[1] Pontificia Univ Catolica Chile, Fac Ciencias Biol, FONDAP Ctr Genome Regulat, Dept Mol Genet & Microbiol, Santiago 8331150, Chile
[2] Pontificia Univ Catolica Chile, Fac Ciencias Biol, Millennium Nucleus Ctr Plant Funct Gen, Dept Mol Genet & Microbiol, Santiago 8331150, Chile
[3] INRA, CNRS, SupAgro UM2,AgroM, Inst Biol Integrat Plantes,Unite Mixte Rech 5004, F-34060 Montpellier 2, France