[1] Univ St Andrews, Sch Psychol & Neurosci, St Andrews, Fife, Scotland
[2] Max Planck Inst Evolutionary Anthropol, Dept Comparat & Dev Psychol, Leipzig, Germany
[3] Univ CEU Cardenal Herrera, Ethol & Anim Welf Sect, Valencia, Spain
[4] CNR, Unit Cognit Primatol, Rome, Italy
[5] CNR, Ctr Primate, Ist Sci & Tecnol Cogniz, Rome, Italy