[1] Oak Ridge Natl Lab, Biosci Div, Oak Ridge, TN 37831 USA
[2] Duke Univ, Dept Biol, Durham, NC 27708 USA
[3] Lorraine Univ, Lab Excellence ARBRE, INRA, UMR 1136, F-54280 Nancy, Champenoux, France
[4] Univ Western Sydney, Hawkesbury Inst Environm, Richmond, NSW, Australia