[1] Univ Limoges, GRESE EA 4330, F-87060 Limoges, France
[2] Univ Auvergne, PIAF UMR547, F-63000 Clermont Ferrand, France
[3] INRA, PIAF UMR547, F-63100 Clermont Ferrand, France
[4] Univ Auvergne, Lab Physiol & Biotechnol Vegetale, F-15013 Aurillac, France
[5] HelmholtzZentrum Munchen, Inst Soil Ecol, D-85764 Neuherberg, Germany
[6] NovaBiom, Ferme Vauventriers, F-28300 Champhol, France