[1] INSERM, U271, Lab Virus Hepat & Pathol Associees, F-69008 Lyon, France
[2] ENS Lyon, INSERM, U412, Lyon, France
[3] Univ Tours, INSERM, ESPRI 3856, Tours, France
[4] Univ Oxford, Glycobiol Inst, Oxford, England
[5] Inst Biol Lille, CNRS, UPR2511, Lille, France