[1] Chinese Acad Sci, Inst Genet & Dev Biol, Ctr Agr Resources Res, Shijiazhuang 050021, Peoples R China
[2] Chinese Acad Sci, Inst Soil & Water Conservat, State Key Lab Soil Eros & Dryland Farming Loess P, Yangling 712100, Peoples R China
[3] Binzhou Univ, Shandong Key Lab Ecoenvironm Sci Yellow River Del, Binzhou 256603, Peoples R China
[4] Qingdao Univ Sci & Technol, Inst Life Sci, Qingdao 266042, Peoples R China
[5] Chinese Acad Sci, Grad Univ, Beijing 10049, Peoples R China