[1] Univ Birmingham, Dept Engn, Birmingham, W Midlands, England
[2] Univ Bergen, Dept Biol Sci, Bergen, Norway
[3] Inst Marine Res, Bergen, Norway
[4] IIASA, Laxenburg, Austria
[5] Marine Scotland, Freshwater Fisheries Lab, Pitlochry, Scotland
[6] Hokkaido Univ, Res Fac Agr, Sapporo, Hokkaido, Japan