[1] Smithsonian Conservat Biol Inst, Conservat Ecol Ctr, Front Royal, VA 22630 USA
[2] Smithsonian Trop Res Inst, Forest Global Earth Observ, Ctr Trop Forest Sci, Panama City, Panama
[3] Pacific Northwest Natl Lab, Joint Global Change Res Inst, College Pk, MD 20740 USA
[4] Univ Illinois, Carl Woese Inst Genom Biol, Urbana, IL 61801 USA
[5] Univ Sheffield, Dept Anim & Plant Sci, Sheffield S10 2TN, S Yorkshire, England
[6] Univ Montana, WA Franke Coll Forestry & Conservat, Bozeman, MT 59812 USA
[7] Univ Montana, Div Biol Sci, Bozeman, MT 59812 USA