[1] Univ Tunis El Manar, Fac Sci Tunis, Dept Life Sci, Biodivers Biotechnol & Climate Change Lab LR11ES0, Tunis 2092, Tunisia
[2] Univ Gafsa, Fac Sci Gafsa, Dept Life Sci, Univ Campus Sidi Ahmed Zarroug, Gafsa 2112, Tunisia
[3] Univ Monastir, Fac Pharm, Lab Anal Treatment & Valorisat Pollutants Environ, Monastir 5000, Tunisia
[4] Univ Gabes, Higher Inst Appl Biol Medenine, Dept Biol Engn, Res Unit Act Biomol Valorisat, Gabes 6029, Tunisia
[5] Ctr Biotechnol Borj Cedria, Lab PlantesExtremophiles, BP 901, Hammam Lif 2050, Tunisia
[6] Inst Murciano Invest & Desarrollo Agr & Medioambi, Dept Biotecnol Genom & Mejora Vegetal, Murcia 30150, Spain
[7] Inst Murciano Invest Biosanitaria IMIB Arrixaca, Murcia 30120, Spain
[8] Inst Murciano Invest & Desarrollo Agr & Medioambi, Dept Desarrollo Rural Enol & Agr Sostenible, Murcia 30150, Spain