[1] Kagoshima Univ, Educ & Res Ctr Marine Resources & Environm, Kagoshima 8900056, Japan
[2] Kyushu Univ, Div Marine Biol Chem, Lab Marine Environm Sci, Dept Biosci & Biotechnol,Fac Agr,Higashi Ku, Fukuoka 8128581, Japan
[3] Natl Res Inst Fisheries & Environm Inland Sea, Environm Conservat Div, Hiroshima 7390452, Japan