[1] Harvard Univ, Dept Earth & Planetary Sci, Cambridge, MA 02138 USA
[2] MIT, Dept Earth Atmospher & Planetary Sci, Cambridge, MA 02139 USA
[3] Max Planck Inst Biogeochem, D-07745 Jena, Germany
[4] Univ Bremen, Ctr Marine Environm Sci MARUM, D-28359 Bremen, Germany
[5] Univ Durham, Dept Earth Sci, Durham DH1 3LE, England