[1] Fed Univ Vicosa UFV, Av Peter Henry Rolfs S-N, BR-36571000 Vicosa, MG, Brazil
[2] Fed Univ Vicosa UFV, Dept Gen Biol DBG, Av Peter Henry Rolfs S-N, BR-36571000 Vicosa, MG, Brazil
[3] Sao Paulo State Agrobusiness Technol Agcy APTA SA, Honey Bee Hlth Lab LASA, POB 07, BR-12400970 Pindamonhangaba, SP, Brazil
[4] Fed Rural Univ Semiarid UFERSA, Dept Anim Sci DCAN, BR110 Km 47, Mossor, RN, Brazil