[1] Natl Inst Infect Dis, Dept Bacteriol, Shinjuku Ku, Tokyo 1628640, Japan
[2] Oita Prefectural Inst Hlth & Environm, Dept Bacteriol, Oita 8701117, Japan
[3] Yamaguchi Prefectural Res Inst Hlth, Div Biol Med, Yamaguchi 7530821, Japan
[4] Osaka Prefectural Inst Publ Hlth, Dept Infect Dis, Osaka 5370025, Japan
[5] Kanagawa Prefectural Publ Hlth Lab, Dept Microbiol, Kanagawa 2530087, Japan
[6] Tokyo Metropolitan Inst Publ Hlth, Dept Bacteriol, Tokyo 1690073, Japan
[7] Toyama Inst Hlth, Dept Bacteriol, Toyama 9390363, Japan
[8] Fukushima Inst Publ Hlth, Dept Microbiol, Fukushima 9608560, Japan