[1] Univ Manchester, Fac Biol Med & Hlth, Sch Biol Sci, Ctr Musculoskeletal Res, Manchester, Lancs, England
[2] Manchester Univ Fdn Trust, Manchester Acad Hlth Sci Ctr, NIHR Manchester Biomed Res Ctr, Manchester, Lancs, England
[3] Univ Leeds, Leeds Inst Rheumat & Musculoskeletal Med, Leeds, W Yorkshire, England
[4] Leeds Teaching Hosp NHS Trust, NIHR Leeds Biomed Res Ctr, Leeds, W Yorkshire, England