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Acad Mil Med Sci, Mil Vet Inst, Key Lab Jilin Prov Zoonosis Prevent & Control, 666 Liuying West Rd, Changchun 130122, Jilin Province, Peoples R China Acad Mil Med Sci, Mil Vet Inst, Key Lab Jilin Prov Zoonosis Prevent & Control, 666 Liuying West Rd, Changchun 130122, Jilin Province, Peoples R China