[1] Univ Oxford, John Radcliffe Hosp, Univ Dept Clin Neurol, Oxford OX3 9DU, England
[2] Univ Oxford, Wellcome Trust Ctr Human Genet, Oxford OX3 7BN, England
[3] Univ British Columbia, Div Neurol, Fac Med, Vancouver, BC V6T 2B5, Canada
[4] Univ British Columbia, Dept Med Genet, Vancouver, BC V6T 2B5, Canada