[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Lab Alpine Ecol & Biodivers, Beijing 100101, Peoples R China
[2] Northwest A&F Univ, State Key Lab Soil Eros & Dryland Farming Loess P, Inst Soil & Water Conservat, Yangling 712100, Peoples R China
[3] Chinese Acad Sci, Northwest Inst Plateau Biol, Haibei Alpine Meadow Ecosyst Res Stn, Key Lab Adaptat & Evolut Plateau Biota, Xining 810008, Peoples R China
[4] Chinese Acad Sci, Grad Univ, Beijing 100049, Peoples R China
[5] Chinese Acad Sci, Kunming Inst Bot, Ctr Mt Ecosyst Studies, Kunming 650201, Peoples R China