[1] Univ Edinburgh, MRC Ctr Inflammat Res, Edinburgh, Midlothian, Scotland
[2] Univ Edinburgh, Ctr Multiple Sclerosis Res, Edinburgh, Midlothian, Scotland
[3] Univ Edinburgh, Ctr Immun Infect & Evolut, Edinburgh, Midlothian, Scotland
[4] Univ Manchester, Manchester Collaborat Ctr Inflammat Res, Manchester, Lancs, England
[5] Univ Edinburgh, Royal Dick Sch Vet Studies, Roslin Inst, Edinburgh EH9 1QH, Midlothian, Scotland