[1] Univ Laval, Dept Sci & Foret, Ctr Rech Mat Renouvelables, Quebec City, PQ G1V 0A6, Canada
[2] No Arizona Univ, Sch Forestry, Flagstaff, AZ 86011 USA
[3] Univ British Columbia, Fac Forestry, 2424 Main Mall, Vancouver, BC V6T 1Z4, Canada
[4] Minist Forets Faune & Parcs Quebec, Direct Inventaires Forestiers, Quebec City, PQ G1H 6R1, Canada
[5] Univ Montpellier, AgroParisTech, CIRAD, CNRS,UMR,TETIS,INRAE, Montpellier, France