[1] Univ Oxford, Sch Geog & Environm, Oxford OX1 3QY, England
[2] Univ Fed Alagoas, Inst Biol & Hlth Sci, Ave Lourival Melo Moto S-N, BR-5707290 Maceio, AL, Brazil
[3] Chico Mendes Inst Biodivers Conservat, Costa Corais Environm Protect Area, Rua Samuel Hardman S-N, BR-55578000 Tamandare, PE, Brazil
[4] Univ Aveiro, DBIO, Aveiro, Portugal
[5] Univ Aveiro, CESAM Ctr Environm & Marine Studies, Aveiro, Portugal