[1] Univ Helsinki, Biomedicum, Haartman Inst, FIN-00014 Helsinki, Finland
[2] Univ Helsinki, Biomedicum, Mol Canc Biol Res Program, FIN-00014 Helsinki, Finland
[3] Univ Helsinki, Cent Hosp, Diagnost Lab, FIN-00014 Helsinki, Finland
[4] Univ Dundee, Ninewells Hosp & Med Sch, Biomed Res Ctr, Dundee DD1 9SY, Scotland