[1] Guangdong Ocean Univ, Coll Fishery, Zhanjiang 524088, Guangdong, Peoples R China
[2] Guangdong Prov Key Lab Pathogen Biol & Epidemiol, Zhanjiang 524088, Peoples R China
[3] Guangdong Key Lab Control Dis Aquat Econ Anim, Zhanjiang 524088, Peoples R China
[4] Zhongkai Univ Agr & Engn, Guangzhou 510225, Guangdong, Peoples R China