[1] Univ Leeds, Mol Med Unit, Leeds, W Yorkshire, England
[2] Royal Victorian Eye & Ear Hosp, Ctr Eye Res, Melbourne, Vic 3002, Australia
[3] Flinders Med Ctr, Dept Ophthalmol, Adelaide, SA, Australia
[4] Imperial Coll Med, Fac Med, London, England
[5] Leicester Royal Infirm, Dept Ophthalmol, Leicester, Leics, England
[6] Univ Manchester, Acad Unit Ophthalmol, Manchester, Lancs, England
[7] Univ Utah, Dept Ophthalmol & Visual Sci, Salt Lake City, UT USA
[8] Univ Utah, Program Human Mol Biol & Genet, Salt Lake City, UT USA