[1] Univ Helsinki, Finnish Museum Nat Hist, Helsinki, Finland
[2] Univ Helsinki, Viikki Plant Sci Ctr ViPS, Dept Biosci, Muse Res Grp, Helsinki, Finland
[3] Inst Adv Stud Koszeg iASK, Koszeg, Hungary
[4] Univ Naples Federico II, Dept Agr Sci, Portici, Italy
[5] Univ Colorado, Dept Ecol & Evolutionary Biol, Boulder, CO 80302 USA
[6] Inst Multidisciplinario Biol Vegetal IMBIV CONICET, Cordoba, Argentina
[7] Univ Turin, Dept Agr Forest & Food Sci DISAFA, Plant Genet, Grugliasco, Italy