[1] Univ Liverpool, Dept Earth Ocean & Ecol Sci, Liverpool L69 3GP, Merseyside, England
[2] Macquarie Univ, Dept Biol Sci, N Ryde, NSW 2109, Australia
[3] Univ Edinburgh, Sch Geosci, Edinburgh EH9 3JN, Midlothian, Scotland
[4] Univ Cape Town, Dept Bot, ZA-7701 Rondebosch, South Africa
[5] N Carolina State Univ, Dept Plant Biol, Raleigh, NC 27695 USA
[6] CSIRO Ecosyst Sci, Trop Ecosyst Res Ctr, Winnellie, NT 0822, Australia