Trees in the Upper Treeline Ecotone in the Polar Urals: Centuries-Old Change and Spatial Patterns

Woody vegetation at the upper limit of its growth is a sensitive indicator of climate change. The aim of this study is to provide an analysis of the centuries-old spatiotemporal dynamics of larch trees at the upper limit of their growth (mountain massif Rai-Iz, Polar Urals, Russia). We used a ground-based method of mapping the remnants of trees that grew in the study area and died during the Little Ice Age. Aerial photographs from the 1960s and high-spatial-resolution satellite images from 2015 were used as data sources to define the locations of trees. Maps of the forest-tundra phytocoenochoras (areas of the terrain that are relatively homogeneous for one or more components of vegetation and/or other indicators) were created using a modified method of boundary detection between forest parcels with different stand densities. The proposed method of boundary detection between the main types of phytocoenochoras allowed us to identify a 15% total increase in areas of closed and open forest and areas with sparse tree growth, as well as a decrease in areas of tundra with single trees over these last decades. Using our spatiotemporal analysis of foresttundra demographics over the last 50 years, we found that the number of trees in the ecotone had doubled. However, modern trees have not yet reached the areas occupied by trees in the past.