[1] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, Herbario Luz Maria Villarreal Puga Inst Bot IBUG, Zapopan, Jalisco, Mexico
[2] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Consejo Nacl Ciencia & Tecnol, Lab Nacl Identificac & Caracterizac Vegetal LaniVe, Zapopan, Jalisco, Mexico
[3] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Invest Mexico Univ Guadalajara, CONACyT, Zapopan, Jalisco, Mexico
[4] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Biosistemat Ecol & Manejo Recursos Nat & Agr BEMAR, Zapopan, Jalisco, Mexico