[1] SUNY Stony Brook, Dept Ecol & Evolut, Stony Brook, NY 11794 USA
[2] Univ Houston, Dept Hlth & Human Performance, Houston, TX USA
[3] Univ Houston, Dept Biol & Biochem, Houston, TX USA
[4] SUNY Stony Brook, Inst Adv Computat Sci, Stony Brook, NY 11794 USA