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Univ Lyon 1, Inserm,U1111,CNRS,UMR5308,LabEx ECOFECT Ecoevolut, Ecole Normale Super Lyon,Ctr Int Rech Infectiol, Equipe Oncogenese Retrovirale,Equipe Labellisee, 46 Allee Italie, F-69007 Lyon, France Univ Lyon 1, Inserm,U1111,CNRS,UMR5308,LabEx ECOFECT Ecoevolut, Ecole Normale Super Lyon,Ctr Int Rech Infectiol, Equipe Oncogenese Retrovirale,Equipe Labellisee, 46 Allee Italie, F-69007 Lyon, France