[1] Univ Melbourne, Fac Sci, Sch Ecosyst & Forest Sci, Richmond, Vic 3121, Australia
[2] Pacific Southwest Res Stn, Urban Ecosyst & Social Dynam Program, Davis, CA 95618 USA
[3] Natl Res Council CNR, Inst Agroenvironm & Forest Biol IBAF, I-05010 Porano, Italy
[4] Acad Sci Czech Republ, Vvi, Global Change Res Ctr, Czechglobe, Brno 60300, Czech Republic